When her family is struck by a terrible tragedy, Sarah Nightingale is forced to go into service at Grey Friars House as an under-nursery-maid. Although the situation wasn’t of her making Sarah thrives there, but life for her younger brother Alfie is not as easy. Desperate to avoid being sent to the workhouse, Alfie runs away to London to seek his fortune. But young and inexperienced with the dangerous streets of London, Alfie is tricked and sold to work as a slave on a coal barge. Sarah misses her brother terribly but knows she must find a way to survive if she ever wants them to be reunited.

Elizabeth Roby lives a content and privileged life in London with husband Jonathon and children, Emily and George. But with the outbreak of war, everything changes. Jonathon informs his family that they have to move from their smart London home to the riverside town of Wivenhoe and their new home Harbour House, where Jonathon will do his duty for his country as part of the Admiralty at the shipyards. But Elizabeth is devastated. How will she start a new life in a place she hardly knows, surrounded by strangers?

Eldest sister Frances is about to marry her childhood sweetheart. But with Marcus Danby now a fighter pilot, his fate hanging in the balance. Will he return from war for their happy ever after? Meanwhile youngest daughter Emily is swept off her feet by a dashing young officer, but does she really know him? Brother Daniel has joined up, but not before falling in love with a local kitchen girl Alice. Against the backdrop of war is there hope of a future for them?

In the heart of Dressmakers' Alley, Madame Pauline’s sweatshop is rumoured to be a particularly unpleasant place to work. Filled with seamstresses paid a pittance who turn a blind eye to what darker activities are hidden behind locked doors. Young Winnie Brown is keen to prove her value to the Women Movement and secures a job as a seamstress to investigate the whispers of some unsavoury goings on. Her concerns are soon justified when she hears a desperate cry for help.

When Sally, Beth, Margaret and Rachel meet at a job interview for the wonderful new store in Oxford Street, they have no idea they will become lifelong friends. When all four girls are lucky enough to be selected as sales staff their exciting new adventure begins. Join them as they overcome heartbreak and grief, find love and happiness and remain united in their friendship, whatever life throws at them.