At the heart of Dover's Market Square lies the Castle's Café. A bustling, homely place run by the formidable Nellie Castle and her six children. Since the scandalous birth of her beloved son 10 years before, Marianne, Nellie's eldest daughter, has preferred to stay in the kitchen, hidden away from the town gossips. Overcome with shame, Marianne has never revealed the name of Donny's father - not even to her own mother. But with World War II just around the corner, Marianne's past catches up with her. And suddenly the lives of the Castle family become a lot more complicated.

1940, London. When Cordelia accepts the post of head librarian in Silver Town Library, her mother is more than a little disapproving. The East End has high levels of poverty and illiteracy, and her mother says it’s no place for a woman of her status. But Cordelia is determined to make a difference in these times of strife, and along with her colleagues, Jane and Mavis, she begins to help the local community, making sure everyone knows what the library can offer them. And maybe even a romance will blossom.

Maureen Jackson knew life as a trainee nurse wouldn't be easy, but she didn't expect her hospital to be badly bombed on her first shift. Plus Maureen still has her family and friends in Mulberry Lane to keep her busy - she's needed as much there as she is by her patients. Running the pub on the corner of Mulberry Lane, Peggy Ashley is used to taking in all sorts of waifs and strays. But the arrival of a dashing American captain has got tongues wagging about Mulberry Lane's favourite landlady....

It is 1939, and working-class Evie Bishop has received a scholarship to study mathematics at Oxford when tragedy turns her life upside down. Evie must seek a new future for herself and, inspired to contribute to the war effort, joins the Women’s Auxiliary Air Force as an Ops Room plotter. Posted to a fighter station on the Sussex Coast, Evie befriends two other WAAFs. But the German bombers seem to know more than they should about the base’s operations, and soon Evie, May and Jess are caught up in a world more dangerous than they ever imagined.

Alice Lake has arrived in London from Liverpool to start her training as a District Nurse, but her journey has been far from easy. Her parents think that she should settle down and get married, but she has already had her heart broken once and isn’t about to make the same mistake again. Alice and her best friend, Edith, are based in the East End, but before they’ve even got their smart new uniforms on, war breaks out, and Hitler’s bombs are raining down on London.

When sixteen-year-old Sally is evacuated to the English south coast, she is terrified by what lies ahead of her. All she knows are the sights and sounds of London's East End – but Sally swallows her tears as they leave the familiar landmarks behind, knowing that she has to be a Grown-Up Girl and play mother to her six-year-old brother Ernie. Playing mother is nothing new for Sally – their real mother Florrie, a good-time girl, hasn't even come to the station to wave them off and Ernie, crippled at an early age by polio, is used to depending on his older sister.

London 1940. As the war rages in Europe, Rose Brown steps on board a ship bound for America, hoping to escape the danger. She's frightened, but her widowed mammy and three sisters in Ireland need the money she earns as nanny to the rich Townsend family. But while Rose plays with baby Sarah in the salty spray on deck, disaster strikes, and the ocean liner begins to sink. In the chaos, Rose and the baby are separated from the family.

Cambridgeshire - March 1939. As the clouds of war begin to gather in Europe, the Talbot family of rural Blackberry Farm will be torn apart, just as so many families all over the world will be. Life will never be the same again. Whilst in London, the Salmons family will feel the pain of parting and loss. Brought together by war, the two families become intertwined and, as the outlook looks bleak, they must draw on each other’s strength to fight through the hard times.

Following a humiliating experience involving the man she thought she’d marry, Iris Tredwick signs up to the Wrens in order to escape and - hopefully - find ‘the right sort’ of man to please her mother. After a bumpy start, Iris manages to befriend outspoken Mary and dreamer Sally as they are sent to their first posting - in Orkney. There she meets engineer Rob, whose flirtatious nature both charms and confounds straight-laced Iris. Much more appropriate for her is local doctor Stewart, if only she felt the same spark for him as she does for Rob.

After renowned fashion designer Cressida Westcott loses both her home and her design house in the London Blitz, she has nowhere to go but the family manor house she fled decades ago. Praying that her niece and nephew will be more hospitable than her brother had been, she arrives with nothing but the clothes she stands in, at a loss as to how to rebuild her business while staying in a quaint country village.

London, 1941. The Blitz. When a Royal Navy memo arrives at head office, requesting female recruits to sign up as motorcycle dispatch riders, delivering highly classified orders across the country, three women jump at the chance to sign up for the most dangerous jobs in London.

Lady Joanna Harcourt and her daughter, Sarah, discover their life of luxury at Goodwill House is over. Because with Lord Harcourt away fighting, the Harcourt girls are facing financial ruin. Lady Joanna is terrified of losing their home, but for Sarah, this means her dreams of becoming a doctor are over. Headstrong Sarah isn’t one to quit! War or not, she’s determined to find a way to save her home and follow her dreams - whatever it takes.

Eldest sister Frances is about to marry her childhood sweetheart. But with Marcus Danby now a fighter pilot, his fate hanging in the balance. Will he return from war for their happy ever after? Meanwhile youngest daughter Emily is swept off her feet by a dashing young officer, but does she really know him? Brother Daniel has joined up, but not before falling in love with a local kitchen girl Alice. Against the backdrop of war is there hope of a future for them?

When Hazel is given the chance to parachute into Nazi-occupied France, she seizes the opportunity to do more for the British war effort than file paperwork. Alongside her childhood friend, French-born Rose, she quickly rises up the ranks of the freedom fighters. For Rose, the Resistance is a link to her late husband, and a way to move forward without him. What starts out as helping downed airmen becomes a bigger cause when they meet Sophia, a German escapee and fierce critic of Hitler who is wanted by the Gestapo.

When Sally, Beth, Margaret and Rachel meet at a job interview for the wonderful new store in Oxford Street, they have no idea they will become lifelong friends. When all four girls are lucky enough to be selected as sales staff their exciting new adventure begins. Join them as they overcome heartbreak and grief, find love and happiness and remain united in their friendship, whatever life throws at them.