Eldest sister Frances is about to marry her childhood sweetheart. But with Marcus Danby now a fighter pilot, his fate hanging in the balance. Will he return from war for their happy ever after? Meanwhile youngest daughter Emily is swept off her feet by a dashing young officer, but does she really know him? Brother Daniel has joined up, but not before falling in love with a local kitchen girl Alice. Against the backdrop of war is there hope of a future for them?

Cambridgeshire - March 1939. As the clouds of war begin to gather in Europe, the Talbot family of rural Blackberry Farm will be torn apart, just as so many families all over the world will be. Life will never be the same again. Whilst in London, the Salmons family will feel the pain of parting and loss. Brought together by war, the two families become intertwined and, as the outlook looks bleak, they must draw on each other’s strength to fight through the hard times.

Born in the workhouse and adopted by a former suffragette, Frances Brakespeare was encouraged from an early age to be strong, independent and to pursue a career as a doctor. The tragic loss of the love of her life in the Great War propels Frances to commit to her one true vocation. Rebelling against the unfair treatment of female doctors Frances is dismissed from her London post and things continue to take a turn for the worse when Izzy, her benefactress dies and Frances finds herself homeless.

With the country on the brink of war, single mother Jean embraces her ten-year-old daughter, Valerie, before she puts her on a train to Somerset alongside hundreds of other evacuees. Jean’s heart breaks as she vows they’ll be together again soon, knowing it’s a promise she might not be able to keep. Wrenched away from her mother and everything familiar, Valerie arrives in Somerset and nervously waits in the village hall to find a host family.

At the heart of Dover's Market Square lies the Castle's Café. A bustling, homely place run by the formidable Nellie Castle and her six children. Since the scandalous birth of her beloved son 10 years before, Marianne, Nellie's eldest daughter, has preferred to stay in the kitchen, hidden away from the town gossips. Overcome with shame, Marianne has never revealed the name of Donny's father - not even to her own mother. But with World War II just around the corner, Marianne's past catches up with her. And suddenly the lives of the Castle family become a lot more complicated.

Amsterdam, 1941. German forces have invaded Amsterdam. As the tension in the city increases, the Dutch Resistance is born. Tormented by what is happening in his city, Christiaan joins the Resistance. He risks everything to shield the oppressed and fight for the freedom of his beloved nation. His brother—Floris—is ruthless and driven to succeed. Promoted to the Bureau of Jewish Affairs, he becomes instrumental in hunting Jews to transport them to the Nazi death camps.

When her family is struck by a terrible tragedy, Sarah Nightingale is forced to go into service at Grey Friars House as an under-nursery-maid. Although the situation wasn’t of her making Sarah thrives there, but life for her younger brother Alfie is not as easy. Desperate to avoid being sent to the workhouse, Alfie runs away to London to seek his fortune. But young and inexperienced with the dangerous streets of London, Alfie is tricked and sold to work as a slave on a coal barge. Sarah misses her brother terribly but knows she must find a way to survive if she ever wants them to be reunited.

In the heart of Dressmakers' Alley, Madame Pauline’s sweatshop is rumoured to be a particularly unpleasant place to work. Filled with seamstresses paid a pittance who turn a blind eye to what darker activities are hidden behind locked doors. Young Winnie Brown is keen to prove her value to the Women Movement and secures a job as a seamstress to investigate the whispers of some unsavoury goings on. Her concerns are soon justified when she hears a desperate cry for help.

When sixteen-year-old Sally is evacuated to the English south coast, she is terrified by what lies ahead of her. All she knows are the sights and sounds of London's East End – but Sally swallows her tears as they leave the familiar landmarks behind, knowing that she has to be a Grown-Up Girl and play mother to her six-year-old brother Ernie. Playing mother is nothing new for Sally – their real mother Florrie, a good-time girl, hasn't even come to the station to wave them off and Ernie, crippled at an early age by polio, is used to depending on his older sister.

As a small child, Nellie Peace was always dreaming but sensed her mother’s rejection. Abandoned and sent into service at Beaumont House at an early age, Nellie is lost and alone until she meets the unpredictable and reclusive artist Lucas Harrington and falls in love with him. This unlikely association between master and servant is encouraged by Lucas’ gentle-natured Aunt Alice, as Lucas sees something unusual in Nellie and is compelled to paint her. Broken promises lead to inevitable heartbreak, and Nellie flees Beaumont House in disgrace for London.

Alice is nursing an injured hand and a broken heart when she moves to the village of Churchwood at the start of WWII. She is desperate to be independent but worries that her injuries will make that impossible. Kate lives with her family on Brimbles Farm, where her father and brothers treat her no better than a servant. With no mother or sisters, and shunned by the locals, Kate longs for a friend of her own. Naomi is looked up to for owning the best house in the village. But privately, she carries the hurts of childlessness.

Three very different girls sign up as trainee nurses at a big London teaching hospital in 1934. Dora leaves her overcrowded, squalid working-class home for a better life. But has she got what it takes to keep up with other, better-educated girls? And will her hated stepfather ever let her go? Helen is born for the job; her brother is a doctor, her all-powerful mother a hospital trustee. But will Helen’s secret misery be her downfall? An aristocratic rebel, Millie’s carefree attitude will find her up in front of Matron again and again. Will she ever care enough to make a nurse? Or will she go back to the glamorous life she was born to?

Elizabeth Roby lives a content and privileged life in London with husband Jonathon and children, Emily and George. But with the outbreak of war, everything changes. Jonathon informs his family that they have to move from their smart London home to the riverside town of Wivenhoe and their new home Harbour House, where Jonathon will do his duty for his country as part of the Admiralty at the shipyards. But Elizabeth is devastated. How will she start a new life in a place she hardly knows, surrounded by strangers?

London, 1938: The bookstore just doesn’t feel the same to Gertie Bingham ever since the death of her beloved husband Harry. Bingham Books was a dream they shared together, and without Harry, Gertie wonders if it’s time to take her faithful old lab, Hemingway, and retire to the seaside. But fate has other plans for Gertie. In Germany, Hitler is on the rise, and Jewish families are making the heart-wrenching decision to send their children away from the growing turmoil. After a nudge from her dear friend Charles, Gertie decides to take in one of these refugees.

Bristol 1936. Jenny Crawford has resigned herself to a loveless marriage living hand to mouth with their two children. Like many others, husband Roy struggles to find work at the docks but a chance encounter turns the family's fortunes around. Not only does he get regular work but they are also allocated a council house on Coronation Close on the outskirts of the city. Jenny and the children are overjoyed, this is the fresh start she could only ever dream of. But trouble feels never too far away.