Clara Newton is the new Housemother of Shilling Grange Orphanage. Many of the children have been bombed out of their homes and left without families, their lives torn apart by the war, just like Clara’s. Devastated by the loss of her fiancé, a brave American pilot, she is just looking for a place to start again. But the orphans are in desperate need of her help.

It’s 1850, and the Noble family have travelled to the other side of the world to start a new life after scandal drove them from their native England. Pippa Noble is determined to reclaim their honour by making her father’s plan for an outback farm a reality, although her ambition is frowned upon by a society that has very set ideas about a woman’s place. Pippa learns the hard way about the unforgiving nature of the bush, sometimes with devastating consequences.

With the heat of their beloved India far behind them, Evie Davenport and her widowed British Army officer father, are starting a new life in England. But Evie is struggling. With her dearest mother gone, Yorkshire with its cold, damp countryside and strict societal rules makes Evie feel suffocated and alone. Her friendship with Sophie Bellingham, the gently reared daughter of a wealthy rail baron, is Evie’s only comfort. Until the arrival of local cotton mill owner, Alexander Lucas. Newly returned from America, it is expected Alexander will marry and finally make England his home.

Yorkshire Dales, 1850. As a terrible storm rages, Annabelle Wallis is shocked to find a distressed young woman at her cottage door, heavy with child. Moments later a baby girl is born. But by dawn, the mother has vanished, leaving behind the helpless child wrapped only in a silk peacock shawl. When news spreads that Lady Eliza Hartley, sister to a wealthy estate owner, has been found dead, Annabelle realises the terrible secret she has stumbled on. Terrified she’ll be blamed for Eliza’s death, Annabelle flees to the slums of York, where she plans to raise the orphan as her own.

Bristol 1936. Jenny Crawford has resigned herself to a loveless marriage living hand to mouth with their two children. Like many others, husband Roy struggles to find work at the docks but a chance encounter turns the family's fortunes around. Not only does he get regular work but they are also allocated a council house on Coronation Close on the outskirts of the city. Jenny and the children are overjoyed, this is the fresh start she could only ever dream of. But trouble feels never too far away.

Maggie Bailey has not had an easy childhood. Her father, Michael, always too easy with his fists after spending the family’s rent money down the pub. Just sixteen, in love and blinded by promises, Maggie sees marriage to handsome Jack as her great escape. However, she soon finds herself abandoned with a beautiful baby when Jack disappears. Maggie is forced to seek a new life away from the East End of London and finds herself a job at a hotel in Yarmouth.

Sophie Whalen is a young Irish immigrant so desperate to get out of a New York tenement that she answers a mail-order bride ad and agrees to marry a man she knows nothing about. San Francisco widower Martin Hocking proves to be as aloof as he is mesmerizingly handsome. Sophie quickly develops deep affection for Kat, Martin's silent five-year-old daughter, but Martin's odd behavior leaves her with the uneasy feeling that something about her newfound situation isn't right.

1932. After the Great War took both her beloved brother and her fiancé, Violet Speedwell has become a "surplus woman," one of a generation doomed to a life of spinsterhood after the war killed so many young men. Yet Violet cannot reconcile herself to a life spent caring for her grieving, embittered mother. After countless meals of boiled eggs and dry toast, she saves enough to move out of her mother's place and into the town of Winchester, home to one of England's grandest cathedrals.

London, 1935. Ruby is always hungry, but she will go without if it means her young brothers can eat. 1930s Bermondsey might be called the larder of London, with its pie, pickle and jam factories, but for the poor working classes, starvation is often only a heartbeat away. When Ruby's neighbour suggests she ought to go to the Methodist Mission for free food, Ruby knows her father will be furious, but she has no other option.

England, 1937: After a devastating childhood at Blakely Hall Orphanage, 15-year-old Sandra is released. She finds work as a housemaid, finally able to put her past behind her. But the start of World War Two throws the country into turmoil, and her brother, Alf, is sent away to fight, leaving her completely alone. Germany, 1939: Eleven-year-old Frieda is about to board a ship bound for England with her brother, Kurt. But at the last moment, as Frieda stands on the deck crammed with frightened children, she spots her brother jumping off, back to land.

When Sally, Beth, Margaret and Rachel meet at a job interview for the wonderful new store in Oxford Street, they have no idea they will become lifelong friends. When all four girls are lucky enough to be selected as sales staff their exciting new adventure begins. Join them as they overcome heartbreak and grief, find love and happiness and remain united in their friendship, whatever life throws at them.

England, 1943: Home is no longer safe for eight-year-old twins Molly and Jacob. Night after night, wailing bombs and screeching planes skim the rooftops overhead. They cradle each other, shivering in terror, not knowing if they'll live to see dawn. Their mother, Martha, has no choice but to evacuate them to the safety of the countryside. At the train station, Martha bites back tears as she says good-bye to her precious children. Knowing she might never see them again, she gives Jacob a letter, pressing the envelope into his hands and telling him to only read it if they are in danger.

Robinswood, Co. Waterford, 1939. The once grand house is home to two very different families. Despite delusions of grandeur, Lord and Lady Kenefick and their adult children live a life of decayed opulence as the money needed to keep such a large house and grounds ever dwindles. Meanwhile, the Murphy family, Dermot, Isabella and their three almost grown-up girls, live and work on the estate and do their best to keep everything running smoothly.

Present day: Emma Lovett, who has dedicated her career to breathing new life into long-neglected gardens, has just been given the opportunity of a lifetime: to restore the gardens of the famed Highbury House estate, designed in 1907 by her hero Venetia Smith. But as Emma dives deeper into the gardens’ past, she begins to uncover secrets that have long lain hidden.

The year is 1731, and in a small village in Cheshire, England, the Reverend Browne has determined the future of his young daughter, Sarah. Her life will be dedicated to caring for him and his household affairs while he, acting as one of God’s chosen few, saves souls from the devil. Sarah dreams of a different future, one in which she will be happy and independent. As she grows older, she starts to see glimpses of a tantalizing world beyond her reach and longs one day to escape the drudgery of her life.